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Malian soldiers backed by French fighter jets battled Islamist rebels in Timbuktu on Sunday after insurgents used a car bomb as cover to infiltrate the northern desert town, sources said.
The French-led offensive in Mali has pushed a mix of Islamists out of their northern strongholds and remote mountain bases but the militants have hit back with several suicide attacks and guerilla-style raids.
At least one Malian soldier was killed and four injured in Sunday's fighting in the ancient Saharan trading hub 1,000 km (600 miles) north of the capital Bamako, according to a Mali government communique issued on Sunday evening.
It said that 21 Islamist rebels were also killed.
"It started after a suicide car bombing around 2200, that served to distract the military and allow a group of jihadists to infiltrate the city by night," said Mali army Captain Modibo Naman Traore.
Bilal Toure, a member of Timbuktu's crisis committee set up after the town was recaptured from Islamist control in January, said he saw a French plane firing on the rebel positions. He said fighting had died down since nightfall.
"The situation settled down after around 1900 but everyone is still staying indoors," he said.
The attack reflected the challenge of securing Mali as France prepares to reduce its troop presence and hand over to the ill-equipped Malian army and a more than 7,000-strong regional African force.
Mali's defense ministry said on Saturday that two Nigerian soldiers in the regional African force were killed when their convoy struck a mine outside Ansongo, near the Niger border.
France launched its intervention in Mali in January to halt an advance by northern al Qaeda-linked rebels towards Bamako.
President Francois Hollande said on Thursday that France will reduce its troop numbers in Mali to 2,000 by July and to 1,000 by the end of the year, down from 4,000 at present.
The West African former colony is to hold presidential and legislative elections in July - vital steps to stabilizing the gold- and cotton-producer after a military coup a year ago paved the way for the northern rebel takeover.